Did officials miss defensive holding call on Iowa State's game-ending interception vs. Oklahoma?

Iowa State beat Oklahoma on Saturday, but did officials miss a penalty call that would have negated the game-deciding play?

Did officials miss defensive holding call on Iowa State's game-ending interception vs. Oklahoma?

Iowa State scored a shocking victory over No. 18 Oklahoma on Saturday, the second loss for the Sooners in as many weeks.

Matt Campbell and his Cyclones secured the 37-30 win on the Sooners' final offensive play of the game when sophomore quarterback Spencer Rattler — trying to march his team down the field for a potential game-winning score — overthrew his receiver, the ball sailing into the waiting hands of Cyclones defender Isheem Young in the end zone.

The Cyclones' victory wasn't without controversy, however: Many who watched the end of the game pointed out how Iowa State's Greg Eisworth II appeared to hold Sooners receiver Charleston Rambo on the play. Had the officials called Eisworth for pass interference or holding, the interception would have been negated and Oklahoma would have had 1:02 to tie or win the game from inside the red zone or just outside it.

Sooners coach Lincoln Riley tried to plead for a call once the play was under review, but to no avail. The Cyclones took over possession, Oklahoma couldn't force a punt, and the Sooners left Ames, Iowa, with their first loss there in 60 years.

If Riley wants to complain about something, then he might start with his own team. There was the dropped would-be touchdown pass earlier in the fourth quarter; Oklahoma would have been facing a three-point deficit on the final drive instead of a touchdown deficit. Or the defense's continued inability to tackle, as Iowa State rolled to 417 total yards and four touchdowns. Or the nine penalties for 93 yards.

The missed holding call may have been the final nail in Oklahoma's coffin on Saturday, but several were already in place.